Retusa

Retusa
Live Retusa sp.
Retusa leptoeneilema
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Order:	Cephalaspidea
Superfamily:	Bulloidea
Family:	Retusidae
Genus:	Retusa T. Brown, 1827
Type species
Bulla obtusa Montagu, 1803
Synonyms
Clistaxis Cossmann, 1895 Coleophysis P. Fischer, 1883 Cryptaxis Jeffreys, 1883 Cylichnina Monterosato, 1884 Mamilloretusa F. Nordsieck, 1972 Retua [sic] (misspelling) Retusa (Coleophysis) P. Pischer, 1883 Retusa (Cylichnina) Monterosato, 1884 Retusa (Retusa) Brown, 1827 Retusa (Sulcularia) Dall, 1921 (invalid: junior homonym of Sulcularia Rafinesque, 1831; Sulcoretusa is a replacement name) Sulcoretusa J. Q. Burch, 1945 Tornatina (Coleophysis) P. Fischer, 1883 Tornatina (Utriculus) T. Brown, 1844 Tornatina (Coleophysis) P. Fischer, 1883 Utriculus T. Brown, 1844 (Invalid: junior homonym of Utriculus Schumacher, 1817) Utriculus (Tornatina) A. Adams, 1850

Retusa is a genus of very small head-shield sea snails or barrel-bubble snails, marine gastropod mollusks in the family Retusidae.

Species

Species within the genus Retusa include:

• Retusa abyssicola Valdés, 2008
• Retusa acrobeles (R. B. Watson, 1883)
• † Retusa acrochone (Cossmann & Pissarro, 1900)
• Retusa agulhasensis Thiele, 1925
• Retusa amboynensis (R. B. Watson, 1883)
• Retusa amphizosta (Watson, 1886)
• Retusa avenaria (R. B. Watson, 1883)
• Retusa atkinsoni (Tenison-Woods, 1876)
• Retusa canariensis (Nordsieck & Talavera, 1979)
• Retusa carpenteri (Hanley, 1859)
• Retusa cecillii (Philippi, 1844)
• Retusa chrysoma Burn in Burn & Bell, 1974
• Retusa chukchii Chaban, 2008
• Retusa complanata (Watson, 1883)
• Retusa concentrica (A. Adams, 1850)
• Retusa crebrisculpta (Monterosato, 1884)
• Retusa crispula (Watson, 1883)
• Retusa crossei (Bucquoy, Dautzenberg & Dollfus, 1886)
• Retusa delicatula (A. Adams, 1862)
• Retusa desgenettii (Audouin, 1826)
• Retusa diaphana Valdés, 2008
• Retusa elegantissima Habe, 1950
• † Retusa ennucleata (Powell & Bartrum, 1929)
• Retusa famelica Watson, 1883
• Retusa frielei (Dall, 1881)
• Retusa galapagana Dall, 1919
• Retusa golikovi Chaban, 2004
• Retusa insolita Valdés, 2008
• Retusa instabilis Minichev, 1971
• Retusa laevisculpta (Granata-Grillo, 1877)
• Retusa lata Chaban & Chernyshev, 2013
• Retusa lenis Valdés, 2008
• Retusa leptekes (R. B. Watson, 1883)
• Retusa leptoeneilema (Brusina, 1866)
• Retusa leuca (R. B. Watson, 1883)
• Retusa longispirata (Yamakawa, 1911)
• Retusa mammillata (Philippi, 1836)
• Retusa mariei (Dautzenberg, 1889)
• Retusa mayoi (Dall, 1889)
• Retusa minima Yamakawa, 1911
• Retusa minutissima (Monterosato, 1878)
• Retusa montereyensis Smith & Gordon, 1948
• Retusa natalensis Barnard, 1963
• Retusa nicobarica Thiele, 1925
• Retusa nitidula (Lovén, 1846)
• Retusa obtusa (Montagu, 1803)
• Retusa operculata Minichev, 1966
• Retusa opima (Finlay, 1926)
• Retusa orientalis Thiele, 1925
• Retusa oruaensis (Webster, 1908)
• Retusa orycta (R. B. Watson, 1883)
• † Retusa otahuhuensis (Laws, 1950)
• Retusa oviformis Thiele, 1925
• Retusa ovoides (Milaschewitsch, 1916)
• Retusa ovulina Lin, 1991
• Retusa parvula (Jeffreys, 1883)
• Retusa paziana Dall, 1919
• Retusa pelyx Burn in Burn & Bell, 1974
• Retusa perforata Thiele, 1925
• Retusa pervia (Dall, 1889)
• Retusa pharetra Hedley, 1909
• † Retusa pressa Marwick, 1928
• Retusa protumida (Hedley, 1903)
• Retusa pseudoglobosa Nomura, 1939
• Retusa pygmaea (A. Adams, 1850)
• Retusa sakuraii (Habe, 1958)
• Retusa sculpta (Gatliff & Gabriel, 1913)
• † Retusa segnis (Laws, 1940)
• Retusa siberutensis Thiele, 1925
• † Retusa soror (Suter, 1917)
• Retusa sosa Ev. Marcus & Er. Marcus, 1969
• Retusa spatha (R. B. Watson, 1883)
• Retusa striata (Hutton, 1873)
• Retusa succincta (A. Adams, 1862)
• Retusa sulcata (d'Orbigny, 1841)
• Retusa sulcata (Watson, 1883) is a secondary junior homonym of Retusa sulcata (d'Orbigny, 1841). No replacement name has been proposed, but the systematics of these small cephalaspids is very confused, and both species may end up in different genera after a proper re-evaluation of their status.
• Retusa sumatrana Thiele, 1925
• Retusa tarutana Smythe, 1979
• Retusa tenerifensis (Nordsieck & Talavera, 1979)
• Retusa tornata (Watson, 1886)
• Retusa toyamaensis (Habe, 1955)
• Retusa trunca Valdés, 2008
• Retusa truncatula (Bruguière, 1792)
• Retusa turrigera Melvill, 1910
• Retusa umbilicata (Montagu, 1803)
• Retusa variabilis (Milaschewitsch, 1912)
• Retusa waughiana Hedley, 1899
• Retusa xystrum Dall, 1919

Taxa inquirenda

• Retusa bysma Melvill, 1904
• Retusa candidula (Locard, 1892) (taxon inquirendum, name is based on a junior primary homonym.)
• Retusa ennurensis Preston, 1916
• Retusa multiquadrata Oberling, 1970 (taxon inquirendum; needs investigation)
• Retusa obesa (Locard, 1897)
• Retusa pusillina (Locard, 1897)
• Retusa robagliana (P. Fischer, 1869) (taxon inquirendum)
• Retusa semen (Reeve, 1855) (taxon inquirendum)

Species brought into synonymy

• Retusa alayoi (Espinosa & Ortea, 2004): synonym of Colinatys alayoi (Espinosa & Ortea, 2004)
• Retusa anderssoni Strebel, 1908: synonym of Diaphana anderssoni (Strebel, 1908)
• Retusa antarctica Melvill & Standen, 1912: synonym of Diaphana paessleri (Strebel, 1905)
• Retusa apiculata (Tate, 1879): synonym of Tornatina apiculata (Tate, 1879): synonym of Acteocina apiculata (Tate, 1879)
• Retusa bizona (A. Adams, 1850): synonym of Mnestia girardi (Audouin, 1826)
• Retusa caelata (Bush, 1885): synonym of Pyrunculus caelatus (Bush, 1885)
• Retusa carinensis de Gregorio, 1889: synonym of Retusa truncatula (Bruguière, 1792)
• Retusa cumberlandiana Strebel, 1908: synonym of Cylichna cumberlandiana (Strebel, 1908)
• Retusa desgenettesi (Audouin, 1827): synonym of Retusa desgenettii (Audouin, 1826)
• Retusa dilatata Pallary, 1904: synonym of Retusa truncatula (Bruguière, 1792)
• Retusa domita (Dall, 1889): synonym of Cylichnium domitum (Dall, 1908)
• Retusa eumicra (Crosse in Crosse & Fischer, 1865): synonym of Tornatina eumicra (Crosse, 1865): synonym of Acteocina eumicra (Crosse, 1865)
• Retusa fourierii (Audouin, 1826): synonym of Pyrunculus fourierii (Audouin, 1826)
• Retusa frigida Hedley, 1916: synonym of Diaphana paessleri (Strebel, 1905)
• Retusa gaimardi Finlay, 1927: synonym of Acteocina decorata (Pilsbry, 1904)
• Retusa girardi (Audouin, 1826): synonym of Ventomnestia girardi (Audouin, 1826)
• Retusa gordonis Yokoyama, 1927: synonym of Acteocina gordonis (Yokoyama, 1927)
• Retusa inflata Strebel, 1908: synonym of Diaphana inflata (Strebel, 1908)
• Retusa insignis (Pilsbry, 1904): synonym of Decorifer insignis (Pilsbry, 1904)
• Retusa lactea (Jeffreys, 1877): synonym of Diaphana lactea (Jeffreys, 1877)
• Retusa mariateresae Parenzan, 1970: synonym of Bulla striata Bruguière, 1792
• Retusa marshalli Sykes, 1904: synonym of Diaphana marshalli (Sykes, 1904)
• Retusa matusimana Nomura, 1939: synonym of Decorifer matusimanus (Nomura, 1939)
• Retusa obtusata: synonym of Retusa obtusa (Montagu, 1803)
• Retusa pellucida (T. Brown, 1827): synonym of Retusa truncatula (Bruguière, 1792)
• Retusa pertenuis (Mighels, 1843): synonym of Retusa obtusa (Montagu, 1803)
• Retusa pfefferi Strebel, 1908: synonym of Diaphana pfefferi (Strebel, 1908)
• Retusa piriformis Nordsieck, 1972: synonym of Retusa truncatula (Bruguière, 1792)
• Retusa semen Thiele, 1925: synonym of Pyrunculus pyriformis (A. Adams, 1850)
• Retusa semisulcata (Philippi, 1836): synonym of Retusa truncatula (Bruguière, 1792)
• Retusa strigella (Lovén, 1846): synonym of Cylichnina umbilicata (Montagu, 1803): synonym of Retusa umbilicata (Montagu, 1803)
• Retusa subcylindrica (Brown, 1827): synonym of Cylichnina umbilicata (Montagu, 1803): synonym of Retusa umbilicata (Montagu, 1803)
• Retusa tohokuensis Nomura, 1939: synonym of Acteocina tohokuensis (Nomura, 1939)
• Retusa truncatoides Nomura, 1939: synonym of Acteocina truncatoides (Nomura, 1939)
• Retusa turrita Møller, 1842: synonym of Retusa obtusa (Montagu, 1803)

References

• Nordsieck, F. (1972). Die europäischen Meeresschnecken (Opisthobranchia mit Pyramidellidae; Rissoacea). Vom Eismeer bis Kapverden, Mittelmeer und Schwarzes Meer. Gustav Fischer, Stuttgart. XIII + 327 pp.
• Gofas, S.; Le Renard, J.; Bouchet, P. (2001). Mollusca, in: Costello, M.J. et al. (Ed.) (2001). European register of marine species: a check-list of the marine species in Europe and a bibliography of guides to their identification. Collection Patrimoines Naturels, 50: pp. 180–213
• Valdés, Á. (2008). Deep-sea "cephalaspidean" heterobranchs (Gastropoda) from the tropical southwest Pacific. In: Héros, V. et al. (eds) Tropical Deep-Sea Benthos 25. Mémoires du Muséum national d'Histoire naturelle. 196: 587-792.

External links

• Brown, T. (1827). Illustrations of the conchology of Great Britain and Ireland. Drawn from nature. W.H. Lizars and D. Lizars, Edinburgh and S. Highley, London. 144 pp., 52 pls
• Brown T. (1844). Illustrations of the Recent Conchology of Great Britain and Ireland, with the description and localities of all the species, marine, land, and fresh water. Ed. 2. London: Smith, Elder & Co. Drawn and Coloured from Nature. Second Edition, Greatly Enlarged
• Burch J.Q. (1945). [Untitled]. Minutes of the Conchological Club of Southern California. 47: 5-26
• Dall, W. H. (1921). Summary of the marine shellbearing mollusks of the northwest coast of America, from San Diego, California, to the Polar Sea, mostly contained in the collection of the United States National Museum, with illustrations of hitherto unfigured species. Bulletin of the United States National Museum. 112: 1-217, pls 1-22.
• Jeffreys, J. G. (1883). On the Mollusca procured during the cruise of H. M. S. Triton, between the Hebrides and Faeroes in 1882. Proceedings of the Zoological Society of London. (1883): 389-399, pl. 44
• Monterosato, T. A. di. (1884). Nomenclatura generica e specifica di alcune conchiglie mediterranee. Virzi, printed for the author, Palermo, 152 pp.
• CZN. (1959). Opinion 568. Protection under the plenary powers of the specific name "obtusa" Montagu, 1803, as published in the combination "Bulla obtusa" (class Gastropoda). Opinions and declarations rendered by the International Commission on Zoological Nomenclature. 20: 404-412.
• Iredale, T. (1915). Some more misused generic names. Proceedings of the Malacological Society of London. 11: 291-306